Curriculum Vitae

RANDY THORNHILL
Department of Biology, MSC03 2020
1 University of New Mexico
Albuquerque, NM 87131-0001

Phone:  505/277-3411
Fax:    505/277-0304
E-mail:   rthorn@unm.edu

Current Rank:   Distinguished Professor, The University of New Mexico

Degrees:

• B.S., Zoology, Auburn University, 1968
  
• M.S., Entomology, Auburn University, 1970
  
• Ph.D., Zoology, The University of Michigan, 1974
  

Ph.D. Dissertation:   Evolutionary Ecology of the Mecoptera (Insecta). 633 pp.

Research Interests:   Evolutionary and ecological aspects of animal social psychology and behavior; insect behavioral ecology; human behavioral ecology and evolutionary psychology; evolutionary methodology.

Research in Progress:

• The evolution of human social psychology and behavior.
• Fluctuating asymmetry and sexual selection.
• Field study of people in a remote Caribbean village.
• The evolution of female sexuality.
• Evolutionary ecology of insects of the order Mecoptera.
• Human values and moral systems in relation to infectious disease.

Recent Ph.D.s:

• Paul Andrews, 2002. Dissertation: The Evolutionary Psychology of Social Chess.
• Kimberly Cline-Brown, 2004. Dissertation: Empirical Evaluation of Evolutionary Hypotheses for Depression. (P.J. Watson, co-chair)
• Corey L. Fincher, 2008. Dissertation: Infectious Diseases and the Ecology and Evolution of Social Life. (P.J. Watson, co-chair)

Current Graduate Students:

• Chris Eppig. Interests:  Sex steroids and dominance in men, concealment of estrus in women, estrus detection by men.
• Kenneth Letendre. Interests:  Psychology of warfare and coalition.

The Human Evolutionary Behavioral Sciences (HEBS) group across the UNM Departments of Anthropology, Biology and Psychology provides faculty for graduate students' dissertation committees.

Teaching Interests:   Undergraduate and graduate level courses in behavioral biology, evolutionary biology, social biology, human evolutionary psychology, and scientific methodology.

Selected Publications:

• In press. S.W. Gangestad, R. Thornhill and C.E. Garver-Apgar. Fertility in the cycle predicts women's interest in attractive bodily features and sexual opportunism. Evolution and Human Bahavior.
• 2009. A.P. Møller, C.L. Fincher and R. Thornhill. Why men have shorter lives than women: Effect of resource availability, infectious diseases, and senescence. American Journal of Human Biology 21:257-364.
• 2009. R. Thornhill, C.L. Fincher and D. Aran. Parasites, democratization, and the liberalization of values across contemporary countries. Biological Reviews 84:113-131.
• 2008. R. Thornhill and S.W. Gangestad. The Evolutionary Biology of Human Female Sexuality. Oxford University Press, New York, NY.
• 2008. P.W. Andrews, S.W. Gangestad, M.G. Haselton, G.F. Miller, R. Thornhill and M.C. Neale. Sex differences in detecting sexual infidelity: Results of a maximum likelihood method for analyzing the sensitivity of sex differences to under-reporting. Human Nature 19:347-373.
• 2008. C.L. Fincher and R. Thornhill. Assortive sociality, limited dispersal, infectious disease, and the genesis of global pattern of religious diversity. Proceedings of the Royal Society of London B 275:2587-2594.
• 2008 C.L. Fincher and R. Thornhill. A parasite-driven wedge: Infectious diseases may explain language and other biodiversity. Oikos 117:1289-1297.
  
• 2008. C.E. Garver-Apgar, S.W. Gangestad and R. Thornhill. Hormonal correlates of women's mid-cycle preference for the scent of symmetry. Evolution and Human Behavior 29:223-232.
• 2008. C.L. Fincher, R. Thornhill, D.R. Murray and M. Schaller. Pathogen prevalence Predicts Human Cross-cultural Variability in Individualism/Collectivism. Proceedings of the Royal Society of London B 275:1279-1285.
• 2008. S.W. Gangestad and R. Thornhill. Human oestrus. Proceedings of the Royal Society of London B 275:991-1000.
• 2007. R. Thornhill and C.L. Fincher. What is the relevance of attachment and life history to political values? Evolution and Human Behavior 28:215-222.
• 2007. R. Thornhill. The importance of developmental biology to evolutionary biology and vice versa. Pp. 203-209 in The Evolution of Mind: Fundamental Questions and Controversies, S.W. Gangestad and J.A. Simpson, eds. Guilford Publications, Inc., New York, NY.
• 2007. R. Thornhill. The evolution of woman’s estrus, extended sexuality and concealed ovulation and their implications for human sexuality research. Pp. 391-396 in The Evolution of Mind: Fundamental Questions and Controversies, S.W. Gangestad and J. A. Simpson, eds. Guilford Publications, Inc., New York, NY.
• 2007. R. Thornhill. Comprehensive knowledge of human evolutionary history requires both adaptationism and phylogenetics. Pp. 31-37 in The Evolution of Mind: Fundamental Questions and Controversies, S.W. Gangestad and J.A. Simpson, eds. Guilford Publications, Inc., New York, NY.
• 2007. S.W. Gangestad and R. Thornhill. The evolution of social inference processes: The importance of signaling theory. Pp. 33-48 in Evolution and the Social Mind, J.P. Forgas, M.G. Haselton and W. von Hippell, eds. Psychology Press, New York, NY
• 2006. C.E. Garver-Apgar, S.W. Gangestad, R. Thornhill, R. Miller and J. Olp. Major histocompatibility complex genes, sexual responsibility, and unfaithfulness in romantic couples. Psychological Science 17:830-835.
• 2006. R. Thornhill. Foreword: Human sperm competition and woman’s dual sexuality. Pp. v-xvii in Sperm Competition in Humans: Classic and Contemporary Readings, T. K. Shackelford and N. Pound, eds. Springer-Verlag Press, New York, NY.
• 2006. R. Thornhill and S.W. Gangestad. 2006. Facial sexual dimorphism, developmental stability and susceptibility to disease in men and women. Evolution and Human Behavior 27:131-144.
• 2006 M. McIntyre, S.W. Gangestad, P.B. Gray, J.F. Chapman, T.C. Burnham, M.T. O’Rourke and R. Thornhill. Romantic involvement often reduces men’s testosterone levels–but not always: The moderating role of extrapair sexual interest. Journal of Personality and Social Psychology 91:642-651.
• 2005. S.W. Gangestad, R. Thornhill and C.E. Garver-Apgar. Adaptations to ovulation: Implications for sexual and social behavior. Current Directions in Psychological Science 14:312-316.
  
• 2005. S.W. Gangestad, R. Thornhill and C.E. Garver-Apgar. Women’s sexual interests across the ovulatory cycle depend on primary partner developmental instability. Proceedings of the Royal Society of London B 272:2023-2027.
• 2005. S.W. Gangestad, R. Thornhill and C.E. Garver-Apgar. Adaptations to ovulation. Pp. 344-371 in The Handbook of Evolutionary Psychology, D.M. Buss, ed. John Wiley and Sons, Hoboken, NJ.
• 2005. A.P. Møller, R. Thornhill and S.W. Gangestad. Direct and indirect tests for publication bias: Asymmetry and sexual selection. Animal Behaviour 70:497-506.
• 2004. R. Thornhill and S.W. Gangestad. The evolution of human attractiveness and attraction. In Evolution: From Molecules to Ecosystems, A. Moya and E. Font, eds. Oxford University Press.
• 2004. S.W. Gangestad and R. Thornhill. Female multiple mating and genetic benefits in humans: Investigations of design. In Sexual Selection in Primates, P. Kappler and C. van Schaik, eds. Cambridge University Press, Cambridge, UK.
• 2004. R. Thornhill and C.T. Palmer. Evolutionary life history perspective on rape. Pp. 249-274 in Evolutionary Psychology, Public Policy and Personal Decisions, C. Crawford and C. Salmon, eds. Lawrence Erlbaum, Mahwah, NJ.
• 2003. C.T. Palmer and R. Thornhill. Straw men and fairy tales: Evaluating reactions to A Natural History of Rape. Journal of Sex Research 40:249-255.
• 2003. R. Thornhill, S.W. Gangestad, R. Miller, G. Scheyd, J. Knight and M. Franklin. MHC, symmetry, and body scent attractiveness in men and women. Behavioral Ecology 14:668-678.
• 2003. R. Thornhill and C.T. Palmer. Rape and evolution: A reply to our critics. Psychology, Evolution and Gender 4:283-296.
• 2003. K. Grammer, B. Fink, A.P. Møller and R. Thornhill. Darwinian aesthetics: Sexual selection and the biology of beauty. Biological Reviews 78:385-407.
• 2003. R. Thornhill and S.W. Gangestad. Evolutionary theory leads to evidence for a male sex pheromone that signals symmetry. Psychological Inquiry 14:316-323.
• 2003. C.T. Palmer and R. Thornhill. A posse of good citizens brings outlaw evolutionists to justice. A response to Evolution, Gender and Rape (2003, ed. by C.B. Travis, MIT Press, Cambridge, MA). Evolutionary Psychology 1:12-27.
• 2003. R. Thornhill and S.W. Gangestad. Do women have evolved adaptation for extra-pair copulation? Pp. 341-368 in Evolutionary Aesthetics, K. Grammer and E. Voland, eds. Springer-Verlag, Berlin, Germany.
• 2003. R. Thornhill. Darwinian aesthetics informs traditional aesthetics. Pp. 9-38 in Evolutionary Aesthetics, K. Grammer and E. Voland, eds. Springer-Verlag, Berlin, Germany.
• 2003. C.T. Palmer and R. Thornhill. Rape and evolution: A reply to our critics. Psychology, Evolution and Gender 4:283-296.
• 2003. S.W. Gangestad and R. Thornhill. Fluctuating asymmetry, developmental stability, and fitness: Toward a model-based interpretation. Pp. 62-80 in Developmental Instability: Causes and Consequences, M. Polak, ed. Oxford University Press, New York.
• 2003. S.W. Gangestad and R. Thornhill. Facial masculinity and fluctuating asymmetry. Evolution and Human Behavior 24:231-241.
• 2003. M. Polak, A.P. Møller, S.W. Gangestad, D.E. Kroeger, J.T. Manning and R. Thornhill. Does an individual asymmetry parameter exist? A meta-analysis. Pp. 81-98 in Developmental Instability: Causes and Consequences, M. Polak, ed. Oxford University Press, New York.
• 2002. Gangestad, S.W., R. Thornhill and C. Garver. 2002. Changes in women’s sexual interest and their partners’ mate retention tactics across the menstrual cycle. Proceedings of the Royal Society of London B 269:975-982.
• 2002. Kowner, R. and R. Thornhill. 2002. The imperfect organism: On the concept of asymmetry and its significance in humans, non-human animals and plants. Symmetry, Culture and Science 10:227-243.
• 2002. K. Grammer, B. Fink, R. Thornhill, A. Juette, and G. Runzal. Female faces and bodies: N-dimensional feature space and attractiveness. In Facial Attractiveness: Evolutionary, Cognitive and Social Perspectives, G. Rhodes and L.A. Zebrowitz, eds. Greenwood Publishers Group, Westport, CT.
• 2001. B. Fink, K. Grammer and R. Thornhill. Human (Homo sapiens) facial attractiveness in relation to skin texture and color. Journal of Comparative Psychology 115:92-99.
• 2001. S. Gangestad, K. Bennett and R. Thornhill. A latent variable model of developmental instability in relation to men’s sexual behavior. Proceedings of the Royal Society of London B 268:1677-1684.
• 2001. R. Thornhill and C.T. Palmer. New preface to paperback edition of A Natural History of Rape: Biological Bases of Sexual Coercion. Http://mitpress.mit.edu/thornhill-preface.html.
• 2001. C.T. Palmer and R. Thornhill. Serial rape: An evolutionary perspective. Pp. 51-65 in Serial Offenders: Current Thought, Recent Findings, Unusual Syndromes, L.B. Schlesinger, ed. CRC Press.
• 2001. C.T. Palmer, R. Thornhill and D.N. Dibari. Biology, sex and the debate over chemical castration. In Sexual Violence: Policies, Practice and Challenges. J.F. Hodgson and D.S. Kelly, eds. Praeger.
• 2000. J.T. Manning, L. Barley, J. Walton, D.I. Lewis–Jones, R.L. Trivers, D. Singh, R. Thornhill, P. Rohde, T. Bereckei, P. Henzi, M. Soler and A. Szwed. The 2^nd:4^th digit ratio, sexual dimorphism, population differences and reproductive success: Evidence for sexually antagonistic genes. Evolution and Human Behavior 21:163-190.
• 2000. R. Thornhill and C.T. Palmer. What A Natural History of Rape really says. The Albuquerque Tribune, March 2, pp. C1-2.
• 2000. R. Thornhill and C.T. Palmer. What A Natural History of Rape really says. The Independent, February 21, p. 5.
• 2000. R. Thornhill and C.T. Palmer. Just why do men rape, authors’ reply. The Sciences, pp. 6, 46-47.
• 2000. R. Thornhill and C.T. Palmer. A Natural History of Rape: Biological Bases of Sexual Coercion. MIT Press.
• 2000. R. Thornhill and C.T. Palmer. Why men rape. The Sciences 40:30-36.
• 1999. R. Thornhill and S.W. Gangestad. Facial attractiveness. Trends in Cognitive Sciences 3:452-460.
• 1999. R. Thornhill. The biology of human rape. Jurimetrics 39:137-147.
• 1999. R. Thornhill. Sexual attraction, the evolutionary psychology of. Pp. 751-753 in The MIT Encyclopedia of the Cognitive Sciences, R.A. Wilson and F.C. Keil, eds. Boston, MA: MIT Press.
• 1999. R. Thornhill and K. Grammer. The body and face of woman: One ornament that signals quality? Evolution and Human Behavior 20:105-120.
• 1999. R. Thornhill and S.W. Gangestad. The scent of symmetry: A human sex pheromone that signals fitness? Evolution and Human Behavior 20:175-201.
• 1999. R. Thornhill, A.P. Møller and S.W. Gangestad. The biological significance of fluctuating asymmetry and sexual selection: A reply to Palmer. American Naturalist 154:234-241.
• 1999. A.P. Møller, S.W. Gangestad and R. Thornhill. Nonlinearity and the importance of fluctuating asymmetry as a predictor of fitness. Oikos 86:366-368.
• 1999. J.E. Scheib, S.W. Gangestad and R. Thornhill. Facial attractiveness, symmetry and cues of good genes. Proceedings of the Royal Society of London, B, 266:1913-1918.
• 1999. J. Manning, R. Trivers, D. Singh and R. Thornhill. The mystery of female beauty. Nature 399:214-215.
• 1999. S.W. Gangestad and R. Thornhill. Individual differences in developmental precision and fluctuating asymmetry: A model and its implications. Journal of Evolutionary Biology 12:402-416.
• 1999. R. Trivers, J. Manning, R. Thornhill, D. Singh and M. McGuire. The Jamaican asymmetry project: Long-term study of fluctuating asymmetry in rural Jamaican children. Human Biology 71:417-430.
• 1998. R. Thornhill and A.P. Møller. The relative importance of size and asymmetry in sexual selection. Behavioral Ecology 9:546-551.
• 1998. G. Arnqvist and R. Thornhill. Evolution of animal genitalia: Patterns of phenotypic and genotypic variation and condition-dependence of genital and non-genital morphology in a water strider (Heteroptera: Gerridae). Genetical Research 71:193-212.
• 1998. A. Møller and R. Thornhill. Male parental care, differential parental investment by females and sexual selection. Animal Behaviour 55:1507-1515.
• 1998. R. Thornhill and F. Bryant Furlow. Stress and human behavior: Attractiveness, women’s sexual development, post-partum depression, and baby’s cry. Pp. 319-369 in Advances in the Study of Behavior, P.J.B. Slater, M. Milinski and A. Møller, eds. Academic Press, New York, NY.
• 1998. S.W. Gangestad and R. Thornhill. Menstrual cycle variation in women’s preferences for the scent of symmetrical men. Proceedings of the Royal Society of London, B, 265:927-933.
• 1998. A. Møller and R. Thornhill. Developmental stability and sexual selection: A meta-analysis. American Naturalist 151:174-192.
• 1998. S.W. Gangestad and R. Thornhill. The analysis of fluctuating asymmetry redux: The robustness of parametric statistics. Animal Behaviour 55:497-501.
• 1998. R. Thornhill. Darwinian aesthetics. Pp. 543-572 in Handbook of Evolutionary Psychology: Ideas, Issues and Applications, C. Crawford and D. Krebs, eds. Lawrence Erlbaum Assoc., Mahwah, NJ.
• 1997. R. Thornhill and A.P. Møller. Developmental stability, disease and medicine. Biological Reviews 72:497-548.
• 1997. R. Thornhill. The concept of an evolved adaptation. Pp. 4-13 in Characterizing Human Psychological Adaptations, G. Bock and G. Cardew, eds. CIBA Foundation, London, UK.
• 1997. G. Arnqvist, R. Thornhill and L. Rowe. Evolution of animal genitalia: Genital and non-genital morphology correlates of fitness components in a water strider. Journal of Evolutionary Biology 10:613-640.
• 1997. A. Møller and R. Thornhill. Developmental instability is heritable: Reply. Journal of Evolutionary Biology 10:69-70.
• 1997. F.B. Furlow, T. Armijo–Prewitt, S.W. Gangestad and R. Thornhill. Psychometric intellegience and fluctuating asymmetry. Proceedings of the Royal Society, London, B, 264.
• 1997. S.W. Gangestad and R. Thornhill. The evolutionary psychology of extra-pair sex: The role of fluctuating asymmetry. Evolution and Human Behavior 18:69-88.
• 1997. S.W. Gangestad and R. Thornhill. Human sexual selection and developmental stability. Pp. 169-195 in Evolutionary Social Psychology, J.A. Simpson and D.T. Kenrick, eds. Lawrence Erlbaum Assoc., Mahwah, NJ.
• 1997. A. Møller and R. Thornhill. A Meta-analysis of the heritability of fluctuating asymmetry. Journal of Evolutionary Biology 10:1-16.
• 1997. R. Thornhill. Rape-victim psychological pain revisited. Pp. 239-249 in Evolution and Human Behavior: A Critical Reader, Laura Betzig, ed. Oxford University Press.
• 1997. J.T. Manning, R.L. Trivers, R. Thornhill, D. Singh, J. Denman, M.H. Eklo, and R.H. Anderton. Ear asymmetry and left-side cradling. Evolution and Human Behavior 18:327-340.
• 1996. R. Thornhill and S.W. Gangestad. Human female copulatory orgasm: A human adaptation or phylogenetic legacy? Animal Behaviour 52:853-855.
• 1996. R. Thornhill and S.W. Gangestad. The evolution of human sexuality. Trends in Ecology and Evolution 11:98-102.
• 1996. B. Furlow and R. Thornhill, The orgasm wars. Psychology Today 29:42-46.
• 1995. A.P. Møller, M. Soler and R. Thornhill. Breast asymmetry, sexual selection and human reproductive success. Ethology and Sociobiology 16:207-219.
• 1995. R. Thornhill, S.W. Gangestad and R. Comer. Human female orgasm and mate fluctuating asymmetry. Animal Behaviour 50:1601-1615.
• 1994. R. Thornhill and S.W. Gangestad. Fluctuating asymmetry correlates with lifetime sex partner numbers and age at first sex in Homo sapiens. Psychological Science 5:297-302.
• 1994. S. Gangestad, R. Thornhill and R. Yeo. Facial physical attractiveness, developmental stability and fluctuating asymmetry. Ethology and Sociobiology 15:73-85.
• 1994. R. Thornhill. Is there psychological adaptation to rape? Analyse and Kritik 16:68-85.
• 1994. P. Watson and R. Thornhill. Fluctuating asymmetry and sexual selection. Trends in Ecology and Evolution 9:21-24.
• 1994. K. Grammer and R. Thornhill. Human (Homo sapiens) facial attractiveness and sexual selection: The role of symmetry and averageness. Journal of Comparative Psychology 108:233-242.
• 1993. R. Thornhill and S. Gangestad. Human facial beauty: Averageness, symmetry and parasite resistance. Human Nature 4:237-269.
• 1993. K. Johnson, R. Thornhill, J.D. Ligon and M. Zuk. The directions of mothers’ and daughters’ preferences and the heritability of male ornaments in red jungle fowl (Gallus gallus). Behavioral Ecology 4:254-259.
• 1993. R. Thornhill. The allure of symmetry. Natural History 102:30-37.
• 1992. M. Zuk, J.D. Ligon and R. Thornhill. Effects of experimental manipulation on male secondary sex characters on female mate preference in red jungle fowl. Animal Behaviour 44:999-1006.
• 1992. R. Thornhill. Fluctuating asymmetry and the mating system of the Japanese scorpionfly, Panorpa japonica. Animal Behaviour 44:867-879.
• 1992. R. Thornhill and N.W. Thornhill. The study of men’s coercive sexuality: What course should it take? Behavioral and Brain Sciences (authors’ response to 28 commentators) 15:404-421.
• 1992. R. Thornhill and N.W. Thornhill. The evolutionary psychology of men’s sexual coercion. Behavioral and Brain Sciences (target article) 15:363-375.
• 1992. R. Thornhill. Fluctuating asymmetry and interspecific competition in two species of Japanese scorpionflies. Behavioral Ecology and Sociobiology 30:357-363.
• 1992. R. Thornhill. Females prefer the pheromone of males with low fluctuating asymmetry in the Japanese scorpionfly (Panorpa japonica). Behavioral Ecology 3:277-283.
• 1992. R. Thornhill and K.P. Sauer. Paternal genetic effects on the fighting ability of sons and daughters and mating success of sons in a scorpionlfy (Panorpa vulgaris). Animal Behavior 43:255-264.
• 1991. S. Austad and R. Thornhill. This bug’s for you. Natural History (No. 12.):44-49.
• 1991. R. Thornhill. Teleonomy and the study of sexual selection. Acta XX Congressus Internationalis Ornitholgici, Vol. III:1361-1366.
• 1991. N.W. Thornhill and R. Thornhill. Evolutionary analysis of psychological pain following rape IV: The effect of the nature of the sex act. J. Comp. Psychology 105:243-252.
• 1991. R. Thornhill and K.P. Sauer. The notal organ of the scorpionfly (Panorpa vulgaris): An adaptation to coerce mating duration. Behavioral Ecology 2:156-164.
• 1990. M. Zuk, R. Thornhill, J.D. Ligon and K. Johnson. Parasites and mate choice in red jungle fowl. American Zoologist 30:235-244.
• 1990. R. Thornhill and N.W. Thornhill. Coercive sexuality of men: Is there psychological adaptation to rape? In Sexual Coercion: Its Nature, Causes and Prevention, E.Grauerholz and M. Koralewski, eds. Lexington Books.
• 1990. M. Zuk, K. Johnson, R. Thornhill and J.D. Ligon. Female red jungle fowl show a threshold mate preference. Evolution 44:477-485.
• 1990. J.D. Ligon, R. Thornhill, M. Zuk, and K. Johnson. Male–male competition and male ornamentation in red jungle fowl. Animal Behavior 40:367-373.
• 1990. R. Thornhill. The study of adaptation. Pp. 31-62 in Interpretation and Explanation in the Study of Behavior, Vol. II, M. Bekoff and D. Jamieson, eds. Westview Press.
• 1990. N.W. Thornhill and R. Thornhill. Evolutionary analysis of psychological pain of rape victims I: The effects of victim’s age and marital status. Ethology and Sociobiology 11:155-176.
• 1990. N.W. Thornhill and R. Thornhill. Evolutionary analysis of psychological pain following rape II: The effects of stranger, friend and family member offenders. Ethology and Sociobiology 11:177-193.
• 1990. N.W. Thornhill and R. Thornhill. Evolutionary analysis of psychological pain following rape III: The effects of force and violence. Aggressive Behavior 16:297-320.
• 1990. M. Zuk, K. Johnson, R. Thornhill and J.D. Ligon. Parasites and sexual selection in red jungle fowl. Behaviour 114:232-248.
• 1990. M. Zuk, R. Thornhill, J.D. Ligon, K. Johnson, S. Ligon, N.W. Thornhill, S. Austad and C. Costin. Female choice and male ornamentation in red jungle fowl. American Naturalist 136:459-473.
• 1989. R. Thornhill and N.W. Thornhill. The evolution of psychological pain. Pp. 73-103 in Sociobiology and the Social Sciences, R. Bell, ed. Texas Tech Univ. Press.
• 1989. R. Thornhill. Nest defense by red jungle fowl (Gallus gallus spadiceus) hens: The roles of renesting potential, parental experience and brood reproductive value. Ethology 83:31-42.
• 1989. R. Montgomerie and R. Thornhill. Fertility advertisement in birds: A means of inciting male–male competition. Ethology 81:209-220.
• 1988. R. Thornhill. The hen’s cackle incites male competition. Veh. Dtsch. Zool. Ges. 81:145-154.
• 1987. N.W. Thornhill and R. Thornhill. Female primate sexual behavior and conception: Are there really sperm to spare? Current Anthropology 29:93-94.
• 1987. L. Heisler, M.B. Andersson, S.J. Arnold, C.R. Boake, G. Borgia, G. Hausfater, M. Kirkpatrick, R. Lande, J. Maynard Smith, P. O’Donald, R. Thornhill and F.J. Weissing. The evolution of mating preferences and attractive traits. In Sexual Selection: Testing the Alternatives, J. Bradbury and M. Andersson, eds. Springer–Verlag, New York, NY.
• 1987. N.W. Thornhill and R. Thornhill. Evolutionary theory and rules of mating and marriage pertaining to relatives. Pp. 373-400 in Psychology and Sociobiology: Ideas, Issues, and Applications, C. Crawford, M. Smith, and D. Krebs, eds. Erlbaum Assoc. Publ., London.
• 1987. R. Thornhill and N.W. Thornhill. Human rape: the strengths of the evolutionary perspective. Pp. 269-291 in Psychology and Sociobiology: Ideas, Issues and Applications, C. Crawford, M. Smith and D. Krebs, eds. Erlbaum Assoc. Publ., London.
• 1987. R. Thornhill. The relative importance of intra- and interspecific competition in scorpionfly mating systems. Am. Nat. 130:711-729.
• 1986. R. Thornhill, N.W. Thornhill and G. Dizinno. The biology of rape. In Rape, S. Tomaseli and R. Porter, eds. Oxford, England: Basil Blackwell and Welcome Institute for the History of Medicine.
• 1986. S. Austad and R. Thornhill. Female reproductive variation in a nuptial-feeding spider, Pisaura mirabilis. Bull. Br. Arachnol. Soc. 7:48-52.
• 1986. R. Thornhill and D. Gwynne. The evolution of sexual differences in insects. Amer. Scientist 74:382-389.
• 1986. R. Thornhill. Relative parental contribution of the sexes to their offspring and the operation of sexual selection. Pp. 113-136 in Evolution of Animal Behavior, M. Nitecki and J. Kitchell, eds. Oxford: Oxford Univ. Press.
• 1985. N.W. Thornhill and R. Thornhill. Matriliny and sexual selection and conflict. Behav. and Brain Sciences 8:679-680.
• 1984. R. Thornhill. Alternative hypotheses for traits believed to have evolved by sperm competition. Pp. 151-178 in Sperm Competition and the Evolution of Animal Mating Systems, R.L. Smith, ed. Academic Press, New York, NY.
• 1984. R. Thornhill. Alternative female choice tactics in the scorpionfly Hylobittacus apicalis (Mecoptera) and its implications. Amer. Zoologist. 24:367-383.
• 1984. R. Thornhill. Fighting and assessment in Harpobittacus scorpionflies (Insecta: Mecoptera). Evolution 38:204-214.
• 1984. R. Thornhill. Scientific methodology in entomology. Florida Entomol. 67:74-96.
• 1983. R. Thornhill. Cryptic female choice in the scorpionfly Harpobittacus nigriceps and its implications. American Naturalist 122:765-788.
• 1983. R. Thornhill and N.W. Thornhill. Human rape: An evolutionary analysis. Ethology and Sociobiology 4:137-173.
• 1983. R. Thornhill, G. Dodson and L. Marshall. Sexual selection and insect mating behavior. American Biology Teacher 45:310-319.
• 1983. R. Thornhill and J. Alcock. The Evolution of Insect Mating Systems. Harvard University Press, Cambridge. 576 pp.
• 1983. G. Byers and R. Thornhill. Biology of the Mecoptera. Ann. Review Entomology 28:203-228.
• 1982. W. Shields, L. Shields, R. Thornhill and N.W. Thornhill. What causes rape? A dissenting view. Science 82 (Dec.):16.
• 1981. R. Thornhill. Panorpa (Mecoptera: Panorpidae) scorpionflies: Systems for understanding resource-defense polygyny and alternative male reproductive efforts. Ann. Rev. Ecol. and Syst. 12:355-386.
• 1980. R. Thornhill. Sexual selection in the black-tipped hangingfly. Scientific American 242:162-172.
• 1980. R. Thornhill. Competitive, charming males and choosy females: Was Darwin correct? Florida Entomologist 63:5-30.
• 1980. R. Thornhill. Mate choice in Hylobittacus apicalis (Insecta: Mecoptera) and its relation to some models of female choice. Evolution 34:519-538.
• 1980. R. Thornhill. Sexual selection within mating swarms of the lovebug, Plecia nearctica (Diptera: Bibionidae). Animal Behavior 28:405-412.
• 1980. R. Thornhill. Rape in Panorpa scorpionflies and a general rape hypothesis. Animal Behavior 28:52-59.
• 1980. R. Thornhill. Competition and coexistence in Panorpa scorpionflies (Mecoptera: Panorpidae). Ecological Monographs 50:179-197.

I am interested in increasing the public’s understanding of science in general and evolutionary biology in particular. In this regard, my research has been featured in televised productions by The Today Show; The Morning Show; CNN Live; CNN International; BBC; National Geographic; Beyond 2000; Discover Magazine; Discover Canada; Dateline NBC; ABC News Special; Charles Grodin Show; Cronkite, Ward & Company’s documentary series; 20/20; Primetime Live; and others, including TV productions in Europe, Australia and South America. Also, my research has been featured in many U.S. and European radio programs, most international newspapers and many international magazines (e.g., Science, Nature, Newsweek, U.S. News and World Report, Economist, Vogue, Playboy, New Scientist, Reader’s Digest, Mademoiselle, Elle, Men’s Health, Health, Time) and in many major newspapers and magazines in the U.S. and many other countries.

Updated July 2009
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